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Asplenium adiantum-nigrum

L. (1753)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Polypodiales Family: Aspleniaceae Genus: Asplenium

‘iwa‘iwa [iwaiwa]

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Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Indigenous

Island Status

Kaua'i Indigenous
O'ahu Indigenous
Molokai Indigenous
Lana'i Indigenous
Maui Indigenous
Hawai'i Indigenous

Dispersal Agents


Bibliography

Name Published In: Sp. Pl.: 1081 (1753)

Other References

Palmer 2003:55 (K, O, Mo, L, M, H); Munro 563 BISH (L†, 1930)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Asplenium adiantum-nigrum East Maui, Kula. On stone wall in Mrs. Walker's front yard. Preserved_Specimen Lyon, H.L. Maui BISH 1926-10-24
2 Asplenium adiantum-nigrum unknown Preserved_Specimen Mann, H. 259 BISH 1864-01-01
3 Asplenium adiantum-nigrum Waimea Canyon State Park, mile 11.3 on Hwy. 550 canyon in remnant forest on steep, dry slope; open, grassy with Pteridium, Pteris and Lythrum Preserved_Specimen Flynn, T.W. 2200 Kauai BISH 1987-05-21
4 Asplenium adiantum-nigrum Waimea District. Makaha Valley, north facing slopes. Diospyros-Acacia-Metrosideros mesic forest w/ Alpitonia ponderosa, Dodonaea viscosa, Nestegis sandwicensis, Psydrax odorata, Chrysodracon aurea, Psychotria mariniana, Kadua affinis, Coprosma waimeae, Myrsine lanaiensis, Polyscias waimeae, Ant Preserved_Specimen Wood, K.R. 17894 Kauai BISH 2018-07-19
5 Asplenium adiantum-nigrum Road to Kona, 5 mi SW of Namakani Paio rock crevices in small kipuka in aa lava Preserved_Specimen Fosberg, F.R. 57491 Hawaii BISH 1978-04-16
6 Asplenium adiantum-nigrum National Park, upper Kealakomo aa boulders under Metrosideros-Dodonaea parkland Preserved_Specimen Stone, B.C. 2959 Hawaii BISH 1959-06-25
7 Asplenium adiantum-nigrum North of Puu Kamaohanui ridge Preserved_Specimen Degener, O. 11416 Oahu BISH 1937-04-17
8 Asplenium adiantum-nigrum E slope of Mauna Loa aa flow Preserved_Specimen Skottsberg, C.J.F. 513 Hawaii BISH 1922-09-15
9 Asplenium adiantum-nigrum Kapulei, E of Kaholoapele, Kamalo ridge, arid rocky slope Preserved_Specimen Degener, O. 3990 Molokai BISH 1928-06-25
10 Asplenium adiantum-nigrum Haleakala, Paliku Very damp Preserved_Specimen Olson, G.E. 97 Maui BISH 1937-08-22
11 Asplenium adiantum-nigrum Oahu dry soil. Preserved_Specimen Judd, J.R. Oahu BISH 1888-01-01
12 Asplenium adiantum-nigrum East Maui, Makawao District, Ulupalakua, near dry Preserved_Specimen Degener, O. 5902 Maui BISH 1927-08-07
13 Asplenium adiantum-nigrum Kona, Keauhou #1 ohia, mamane grassland Preserved_Specimen Christ, J.H. Hawaii BISH 1958-05-15
14 Asplenium adiantum-nigrum Puu Hualalai, Puuwaawaa edge of moist gulch Preserved_Specimen St.John, H. 11409 Hawaii BISH 1931-12-29
15 Asplenium adiantum-nigrum 2nd gulch S. of Pohakea pass, Waianae Mts., Halona, Lualualei Forest Reserve steep, grassy slope Preserved_Specimen St.John, H. 13148 Oahu BISH 1933-05-12
16 Asplenium adiantum-nigrum Haleakala, ~5 miles from entrance side of road Preserved_Specimen Tanabe, L. 59 Maui BISH 1961-01-02
17 Asplenium adiantum-nigrum Waianae Mts., Puu Manawahua grassy W. slope Preserved_Specimen St.John, H. 10113 Oahu BISH 1929-12-08
18 Asplenium adiantum-nigrum Saddle Road deep in lava cracks in lava field, descending towards wet side Preserved_Specimen van der Werff, H. Hawaii BISH 1973-09-25
19 Asplenium adiantum-nigrum Haleakala Crater rim, near Rest House rocky slope Preserved_Specimen Fosberg, F.R. 9962 Maui BISH 1933-08-21
20 Asplenium adiantum-nigrum SW flank of Mauna Loa, Manuka Open Metrosideros on cinder-scrub vegetation, Machaerina, Coprosma, Styphelia, Vaccinium Preserved_Specimen Higashino, P.K. 6216 Hawaii BISH 1977-07-21
21 Asplenium adiantum-nigrum East Maui, Kaup Gap in gulch east of trail in gulch Preserved_Specimen Welton, P. 2373 Maui BISH 2016-09-06
22 Asplenium adiantum-nigrum Crest and E. side of Makolelau ridge from Puu Kolekole heading S. along old road to coast dry forest of Diospyros, Pouteria, Canthium, Nestegis, Pleomele; in shade under tree, dry gulch Preserved_Specimen Lorence, D.H. 5665 Molokai BISH 1987-10-23
23 Asplenium adiantum-nigrum near Koolau Gap, within Haleakala Crater moderately dry Preserved_Specimen Degener, O. 3988 Maui BISH 1927-04-15
24 Asplenium adiantum-nigrum Saddle Road, 6 3/4 miles E of Puu Huluhulu Mossy woods of Metrosideros in kipuka. Preserved_Specimen St.John, H. 25374 Hawaii BISH 1954-12-20
25 Asplenium adiantum-nigrum Puu Manawahua rocks Preserved_Specimen Topping, D.L. 3606 Oahu BISH 1932-09-05
26 Asplenium adiantum-nigrum East Maui, Makawao District, Halemauu trail, near Haleakala road crevice of lava rock Preserved_Specimen Wilson, K.A. 62 Maui BISH 1951-12-24
27 Asplenium adiantum-nigrum 23 mile marker, Saddle Road crevices in mostly barren lava substrate Preserved_Specimen Takeuchi, W.N. 625 Hawaii BISH 1983-11-26
28 Asplenium adiantum-nigrum N. Hilo, along Saddle Road near mile marker 23, W of Hilo barren aa flow Preserved_Specimen Lammers, T.G. 5433 Hawaii BISH 1983-09-04
29 Asplenium adiantum-nigrum Just off powerline road 3 miles from Saddle Road 23 mile marker On ground in grassy area. Preserved_Specimen Waters, B. Hawaii BISH 1994-08-27
30 Asplenium adiantum-nigrum Waimea, upper Kawai Iki ridge mesic forest, 60% closed 10-12 m canopy; Metrosideros, Acacia, Dodonaea viscosa, Santalum freycinetianum, Melicope ovata, M. barbigera, Poa sandwicensis, Gahnia beecheyi Preserved_Specimen Wood, K.R. 11996 Kauai BISH 2005-08-03
31 Asplenium adiantum-nigrum 6 mi up paved road toward Mauna Loa from jct. with Rte. 20; turnoff is just E of Puu Huluhulu Tan pahoehoe lava. Scattered in lava crevices. Preserved_Specimen Stuessy, T.F. 6153 Hawaii BISH 1982-05-30
32 Asplenium adiantum-nigrum Kaupulehu under fall fountain grass; dryland forest dominated by Diospyros and Colubrina Preserved_Specimen Takeuchi, W.N. 3542 Hawaii BISH 1987-05-01
33 Asplenium adiantum-nigrum Pohakea Pass under lantana Preserved_Specimen Degener, O. 10454 Oahu BISH 1936-03-01
34 Asplenium adiantum-nigrum East Maui, Makawao, ~0.5 mi below Kula Lodge along Haleakala Hwy small gulch dominated by pasture grasses Preserved_Specimen Wagner, W.L. 5816 Maui BISH 1988-03-05
35 Asplenium adiantum-nigrum Laumaia just above the timber line Preserved_Specimen Neal, M.C. 856 Hawaii BISH 1935-08-18
36 Asplenium adiantum-nigrum Northern slopes, Puu Palikea. N slopes, dry hillside Preserved_Specimen Topping, D.L. 3613 Oahu BISH 1932-09-17
37 Asplenium adiantum-nigrum Mt. Palikea dry ridge Preserved_Specimen Topping, D.L. 3347 Oahu BISH 1929-07-07
38 Asplenium adiantum-nigrum Mauna Kea crevice Preserved_Specimen Neal, M.C. 791 Hawaii BISH 1935-08-12
39 Asplenium adiantum-nigrum Kilauea dry light woods Preserved_Specimen Degener, O. 186 Hawaii BISH 1922-12-13
40 Asplenium adiantum-nigrum National Park, Kipuka Ki dry aa lava Preserved_Specimen Webster, G.L. 1718 Hawaii BISH 1948-06-16
41 Asplenium adiantum-nigrum Puuwaawaa Metrosideros-Grevillea forest on transect T17, aa crevices Preserved_Specimen Takeuchi, W.N. 5834 Hawaii BISH 1989-05-01
42 Asplenium adiantum-nigrum Hanaula, West Maui. Found in thick Dodonaea, Acacia cf. koaie scrub on side of gully. Preserved_Specimen Medeiros, A.C. 534 Maui BISH 1983-09-27
43 Asplenium adiantum-nigrum East Maui, Koolau Forest Reserve gulch with grasses, vaccinium, Rubus, and other ferns Preserved_Specimen Harrison, B. 504 Maui BISH 1973-11-09
44 Asplenium adiantum-nigrum SW flank of Mauna Loa, Manuka, Puu Ohohia closed Metrosideros forest, cinder, understory Deschampsia, Myoporum, Sophora, Styphelia, Vaccinium Preserved_Specimen Higashino, P.K. 6322 Hawaii BISH 1977-08-05
45 Asplenium adiantum-nigrum Manuka NAR;, nature trail up to Kipuka Kilioe Metrosideros lowland mesic forest ca 25 - 30 m tall. W/ understory of Nestegis, Pisonia, Charpentiera, Psychotria hawaiiensis, Claoxylon, Myrsine lessertiana, Dodonaea viscosa, Alyxia oliviformis, Pipturus albidus, Antidesma platyphyllum & A. pulvinatum. Preserved_Specimen Wood, K.R. 13204 Hawaii BISH 2008-08-07
46 Asplenium adiantum-nigrum Pohokea pass, Waianae Mts., Halona, Lualualei Forest Reserve wooded slope Preserved_Specimen St.John, H. 13132 Oahu BISH 1933-05-12
47 Asplenium adiantum-nigrum Pihea Trail across plateau, between Pihea and Alakai Swamp Trail mossy bank in very wet, irregular scrub montane rain forest, or cloud forest Preserved_Specimen Fosberg, F.R. 64356 Kauai BISH 1984-03-31
48 Asplenium adiantum-nigrum Mauna Kea, Wai Kahalulu Gulch adz quarry, shelter cave Preserved_Specimen Stone, F.D. Hawaii BISH 1991-08-10
49 Asplenium adiantum-nigrum E. of Humuula, below Kalaieha pahoehoe Preserved_Specimen Neal, M.C. 704 Hawaii BISH 1935-08-12
50 Asplenium adiantum-nigrum Puu Keekee; 0.5 mi NW, just SE of Parker Ranch fenceline rockpile, Myoporum-Chamaesyce Preserved_Specimen Wagner, W.L. 4684 Hawaii BISH 1982-11-17
51 Asplenium adiantum-nigrum Haleakala, 2 miles from entrance side of road with other mosses and ferns Preserved_Specimen Tanabe, L. 61 Maui BISH 1961-01-02
52 Asplenium adiantum-nigrum Mauna Kea, Parker Ranch Sheep Station dry rocks Preserved_Specimen Topping, D.L. 3628 Hawaii BISH 1932-10-11
53 Asplenium adiantum-nigrum Mauna Loa, Mokuaweoweo 1940 lava flow, warm crack Preserved_Specimen Barrel, R.L. Hawaii BISH 1960-05-19
54 Asplenium adiantum-nigrum Waiamao Hill beyond Waiohinu Preserved_Specimen Giffard, W.M. 42 Hawaii BISH 1927-07-28
55 Asplenium adiantum-nigrum Makawao District. Hanakauhi, Kalialinui subalpine to West of Puu Alaea, TNC Waikamoi Preserve Sophora chysophylla-leptecophyll tameiaemeiae shrubland. With Coprosma montana, Metrosideros spp. Geranium cuneatum, Pteridium decompositum, Sadleria cyathioides, Holcus lantus. On dry stream banks Preserved_Specimen Perlman, S. 23306 Maui BISH 2013-02-28
56 Asplenium adiantum-nigrum Kilauea, Bird Park dry open woods Preserved_Specimen Degener, O. 5905 Hawaii BISH 1929-07-09
57 Asplenium adiantum-nigrum Hilo forest Preserved_Specimen Newell, Brother M. Hawaii BISH 1910-01-01
58 Asplenium adiantum-nigrum below Puu Kolekole ridge Preserved_Specimen Forbes, C.N. 104 Molokai BISH 1912-07-01
59 Asplenium adiantum-nigrum Pohakea pass, Waianae Mts., Honouliuli dry ridge Preserved_Specimen Fosberg, F.R. 9505 Oahu BISH 1933-05-12
60 Asplenium adiantum-nigrum East Maui, Hana District, Hanawi Natural Area Reserve, Hanawi drainage basin, west fork Terrestrial, on densely shaded slope in soil. Metrosideros, Cheirodendron, Coprosma, Melicope, Ilex, Myrisne, Dubautia, Rubus, Sadleria, Vaccinium, Dryopteris, Athyrium, Polystichum, Oerogrammitis, Elaphoglossum, Asplenium. Preserved_Specimen Oppenheimer, H.L. 101113 Maui BISH 2011-10-13
61 Asplenium adiantum-nigrum Hualalai, N. Kona deep cinder cone, water dripping from side; with Polystichum, Metrosideros, Railliardia, Stenogyne, Styphelia Preserved_Specimen Higashino, P.K. 6625 Hawaii BISH 1977-11-26
62 Asplenium adiantum-nigrum Kona, Kealakekua Ranch, near Puu o Uo aa, scattered colonies of vegetation Preserved_Specimen Christ, J.H. Hawaii BISH 1958-11-23
63 Asplenium adiantum-nigrum Kamaileunu. Northern Waianae mountains. On dry open ridge at highest point of Kamaileunu. 3 plants total. Preserved_Specimen Kawelo, K. 591 Oahu BISH 2024-10-09
64 Asplenium adiantum-nigrum Kahuku dry, hot exposed aa flow Preserved_Specimen Russ, G.W. 45 Hawaii BISH 1927-06-01
65 Asplenium adiantum-nigrum EASt Maui, Kaupo Gap in gulch east of trail in gulch Preserved_Specimen Welton, P. 2382 Maui BISH 2017-03-07
66 Asplenium adiantum-nigrum Haleakala, small lava crater S side of Kauaiki dry basalt, in scrub Preserved_Specimen St.John, H. 21227 Maui BISH 1945-08-28
67 Asplenium adiantum-nigrum Waimea Forest. Preserved_Specimen Lyons, A.B. Hawaii BISH 1860-01-01
68 Asplenium adiantum-nigrum Within Haleakala Crater In dry locality Preserved_Specimen Degener, O. 3991 Maui BISH 1927-08-09
69 Asplenium adiantum-nigrum N outer slope of Haleakala, 1.5 mi N of Paliku Cabins heath scrub in shaded low area among Deschampsia Preserved_Specimen Henrickson, J.S. 3670 Maui BISH 1969-06-22
70 Asplenium adiantum-nigrum Puu Kolekole slopes Preserved_Specimen Forbes, C.N. 129 Molokai BISH 1912-07-01
71 Asplenium adiantum-nigrum Waikekeehia, foot of pali E. of Haleakala crater turfy ledge Preserved_Specimen St.John, H. 17746 Maui BISH 1936-12-25
72 Asplenium adiantum-nigrum down Chain of Craters Road ~12.9 mi toward Kalapana from intersection with Crater Rim Rd.; across from turnoff to Naulu Picnic area (plot 5) Metrosideros-Diospyros mixed forest with patchy stands of Aleurites on `a`a lava. Preserved_Specimen Newell, C.L. 477 Hawaii BISH 1966-08-20
73 Asplenium adiantum-nigrum Kauai Preserved_Specimen Brodie, A. 4053 Kauai BISH
74 Asplenium adiantum-nigrum Haleakala Hwy. 20.8N 156.3W Preserved_Specimen Krajina, V.J. 620415051 Maui BISH 1962-04-15
75 Asplenium adiantum-nigrum East Maui Preserved_Specimen Degener, O. 5903 Maui BISH 1927-06-23
76 Asplenium adiantum-nigrum E. of Uhulele Preserved_Specimen Forbes, C.N. 879 Maui BISH 1919-07-21
77 Asplenium adiantum-nigrum Hawaii Preserved_Specimen U.S. Exploring Expedition 43 Hawaii BISH 1838-01-01
78 Asplenium adiantum-nigrum Ulupalakua, Auhi Preserved_Specimen Munro, G.C. 386 Maui BISH 1915-02-12
79 Asplenium adiantum-nigrum Humuula trail, Mauna Kea Preserved_Specimen Hartt, C.E. 760 Hawaii BISH 1935-08-09
80 Asplenium adiantum-nigrum Waiamao Hill, near Waiohinu, Kau Preserved_Specimen Giffard, W.M. 42 Hawaii BISH 1927-07-28
81 Asplenium adiantum-nigrum Waihou Spring, Olinda Preserved_Specimen Hobdy, R.W. 348 Maui BISH 1978-10-23
82 Asplenium adiantum-nigrum Hawaii Preserved_Specimen Mann, H. 259 Hawaii BISH 1864-01-01
83 Asplenium adiantum-nigrum Paliakoae Preserved_Specimen Munro, G.C. 563 Lanai BISH 1930-09-30
84 Asplenium adiantum-nigrum Hawaii Preserved_Specimen Rémy, J. 36 Hawaii BISH 1851-01-01
85 Asplenium adiantum-nigrum Paliakoae (with Waiakeakua scrated out on typed line below). Preserved_Specimen Munro, G.C. 61 Lanai BISH 1927-06-07
86 Asplenium adiantum-nigrum Kaeanae, Halehoku, Haleakala Preserved_Specimen Forbes, C.N. 1008 Maui BISH 1919-08-02
87 Asplenium adiantum-nigrum Between Kula Sanitarium and Polipoli Springs Preserved_Specimen Topping, D.L. 3823 Maui BISH 1935-09-04
88 Asplenium adiantum-nigrum Mauna Loa; E. slopes at road from Kilauea 19.4N 155.3W Preserved_Specimen Krajina, V.J. 611226040 Hawaii BISH 1961-12-26
89 Asplenium adiantum-nigrum Summit of Haleakala Preserved_Specimen Munro, G.C. 653 Maui BISH 1918-04-24
90 Asplenium adiantum-nigrum Ulupalakua, E Maui, Makawao District Preserved_Specimen Munro, G.C. 159 Maui BISH 1915-02-12
91 Asplenium adiantum-nigrum Waimea Drainage Basin, West side Preserved_Specimen Forbes, C.N. 813 Kauai BISH 1917-07-03
92 Asplenium adiantum-nigrum Polipoli Rd., Kula Forest Reserve Preserved_Specimen Hobdy, R.W. 263 Maui BISH 1972-02-29
93 Asplenium adiantum-nigrum Kilauea, Lava Tube Region Preserved_Specimen Morley, H. 75 Hawaii BISH 1934-08-06
94 Asplenium adiantum-nigrum Waimea Drainage Basin, west side. Preserved_Specimen Forbes, C.N. 813 Kauai BISH 1917-07-03
95 Asplenium adiantum-nigrum N mauka of Ulupalakua Preserved_Specimen Degener, O. 3989 Maui BISH 1927-07-04
96 Asplenium adiantum-nigrum E of Ukulele Preserved_Specimen Forbes, C.N. 810 Maui BISH 1919-07-17
97 Asplenium adiantum-nigrum Haleakala Preserved_Specimen Whitney, J.M. Maui BISH 1870-01-01
98 Asplenium adiantum-nigrum Mauna Kea, near Rockshelter 14-R1 Preserved_Specimen McEldowney, H. Hawaii BISH 1976-09-06
99 Asplenium adiantum-nigrum National Park Preserved_Specimen Walker, E. Hawaii BISH 1932-05-06
100 Asplenium adiantum-nigrum Kilauea, Bird Park Preserved_Specimen Neal, M.C. Hawaii BISH 1929-08-05
101 Asplenium adiantum-nigrum Preserved_Specimen Brodie, A. Kauai BISH 1904-07-24
102 Asplenium adiantum-nigrum East Maui, Haleakala Preserved_Specimen Anderson, D. Maui BISH 1932-08-20
103 Asplenium adiantum-nigrum Mauna Loa Trunk road Preserved_Specimen Cranwell, L.M. Hawaii BISH 1938-09-14
104 Asplenium adiantum-nigrum Preserved_Specimen C.W. Seelye BISH
105 Asplenium adiantum-nigrum Puukapele. Preserved_Specimen Brodie, A. Kauai BISH 1904-11-01
106 Asplenium adiantum-nigrum Hualalai Preserved_Specimen Meebold, A. Hawaii BISH 1932-05-01
107 Asplenium adiantum-nigrum Mauna Kea, W of summit Preserved_Specimen Judd, C.S. Hawaii BISH 1935-08-08
108 Asplenium adiantum-nigrum unknown Preserved_Specimen Lack, F. BISH 1880-01-01
109 Asplenium adiantum-nigrum Waikola Preserved_Specimen Hobdy, R.W. 2448 Hawaii BISH 1985-09-12
110 Asplenium adiantum-nigrum Kilauea, Bird Park Preserved_Specimen Smith, F.G. 38 Hawaii BISH 1929-03-14
111 Asplenium adiantum-nigrum Below Puu Oo Preserved_Specimen Forbes, C.N. 812 Hawaii BISH 1915-06-11
112 Asplenium adiantum-nigrum Kilauea, Bird Park Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-08-09
113 Asplenium adiantum-nigrum Haleakala Crater, near Rest House Preserved_Specimen Maui BISH 1935-01-05
114 Asplenium adiantum-nigrum East Maui, crater of Haleakala Preserved_Specimen Forbes, C.N. 303 Maui BISH 1910-08-01
115 Asplenium adiantum-nigrum Waianae Mts., Puu Manawahua. Preserved_Specimen Hosaka, E.Y. Oahu BISH 1929-12-08
116 Asplenium adiantum-nigrum unknown Preserved_Specimen Baldwin, D.D. BISH
117 Asplenium adiantum-nigrum unknown Preserved_Specimen Hillebrand, W.B. BISH
118 Asplenium adiantum-nigrum unknown: Hawaii, Maui, and Kauai. Preserved_Specimen BISH
119 Asplenium adiantum-nigrum unknown Preserved_Specimen BISH
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